|A review of the southern African genus Tongamya (Diptera: Mydidae: Megascelinae), with a molecular assessment of the phylogenetic placement of Tongamya and the Megascelinae|
|Irwin, ME, Wiegmann, BM|
|Afrotropical, Megascelinae, Mydidae, phylogeny, Tongamya|
This paper both tests the morphologically based cladistic appraisal of the Apioceridae proposed by Yeates & Irwin (1996) using the 28S ribosomal RNA gene and reviews the systematic status and biology of Tongamya and the subfamily Megascelinae to which it belongs. Yeates & Irwin (1996) concluded that the subfamilies Rhaphiomidinae and Megascelinae, both formerly placed within the Apioceridae, were separate basal lineages of the family Mydidae. The 28S rDNA evidence presented here confirms the sister group relationship of Tongamya and Megascelus, their retention in the subfamily Megascelinae, the removal of that subfamily from the Apioceridae and its placement within the family Mydidae. New material and field observations of flies belonging to the genus Tongamya enable us to expand the known range of Tongamya miranda, describe a second species, Tongamya stuckenbergi sp. n., better diagnose Tongamya and its two included species, and expand on the habitat and flight behaviour of this remarkable and rarely encountered genus of flies.