|Studies of Mydidae (Diptera). V. Phylogenetic and biogeographic notes, key to the American genera and illustrations of spermathecae|
|Artigas, JN, Papavero, N|
|Anomalomydas, Anomalomydinae, Apiophorinae, Ceriomydas, Diochlistinae, Dolichogaster, Ectyphinae, Gauromydas, Heteromydas, Leptomydinae, Mapinguari, Messiasia, Miltinus, Mitrodetus, Mydas, Mydidae, Mydinae, Nemomydas, Opomydas, Phyllomydas, Protomydas, Pseudonomoneura, Pseudorhopalia, Rhopaliinae|
The Diochlistinae (comprising the genera Mitrodetus Gerstaecker, from Chile, and Diochlistus Gerstaecker and Triclonus Gerstaecker, from Australia) are considered the sister-group of all the remaining Mydidae. The Diochlistinae present as symplesiomorphy separate veins M1 and M2, and as synapomorphies an elongated scape and the gonostyli internally fused to the hypandrium. The remaining Mydidae have as synapomorphy the fusion of veins M1 and M2. A phylogeny of the three genera of Diochlistinae is presented, with Mitrodetus as the sister-group of the genera Diochlistus and Triclonus. It is postulated that the Diochlistinae and their sister-group were sympatrically distributed over Pangaea, and that the Diochlistinae were extinct all over that supercontinent, except in Southern South America and Australia. A key to the American genera of Mydidae is presented, and illustrations furnished of the spermathecae of Mitrodetus, Triclonus, Anomalomydas, Opomydas, Heteromydas, Pseudorhopalia, Miltinus, Dolichogaster, Messiasia, Phyllomydas, Protomydas, Gauromydas, Mydas, Ceriomydas, Mapinguari, Nemomydas, and Pseudonomoneura.