deciphering their diversity and evolutionary history

Smithsonian National Museum of Natural History

Asiloid Flies
deciphering their diversity and evolutionary history

Sample of Asiloid Flies

South Africa trip Sep-Oct 2009

Field trip to western South Africa and visit to the Iziko South African Museum (20 September–5 October 2009)

Funded by U.S. National Science Foundation REVSYS grant (DEB-0919333, local link, NSF link).

Torsten visited the Iziko South African Museum in Cape Town where 106 holotypes of southern African Mydidae and two of the three South African Apiocera are deposited. The images of type specimens that were taken will be of great help for identification of specimens that have been borrowed from numerous museum collections. Specimens for a morphological phylogeny of Apioceridae and Mydidae were also borrowed.

Jason Londt from the Natal Museum in Pietermaritzburg (KwaZulu-Natal, South Africa) joined Torsten for field work in the Western and Northern Cape provinces. We visited quite a few type localities from which Mydidae species flying in the southern Hemisphere spring had been described, but were only able to collect 6 Mydidae species. Unfortunately, we didn't catch any of the more enigmatic genera like Agaperemius, Halterorchis, or Namadytes, which are known from this area.

Numerous Asilidae species that are important for the REVSYS project were collected though. In particular, specimens of Acnephalum and Sisyrnodytes are very important to test the monophyly of the Willistonininae hypothesised by Torsten in 2009 to be a subfamily taxon known only from western North America and southern Africa (Sisyrnodytes does also occur in the Mediterranean).

Below are some images of places that we visited and some additional wildlife:

Habitat next to a dry river bed of the Droe River (near Vanrhynsdorp, Western Cape) where Arenomydas caerulescens (Mydidae: Syllegomydinae) was collected.

Habitat next to a dry river bed of the Droe River (near Vanrhynsdorp, Western Cape) where Arenomydas caerulescens (Mydidae: Syllegomydinae) was collected.

White, vegetated sand dunes 8 km E Port Nolloth (Northern Cape) where a Cephalocera species (Mydidae: Syllegomydinae) was collected.

White, vegetated sand dunes 8 km E Port Nolloth (Northern Cape) where a Cephalocera species (Mydidae: Syllegomydinae) was collected. Note the wall of fog over the cold Benguela current in the distance.

Red, vegetated sand dunes (near Klipheuwel, Western Cape) where Afroleptomydas opacicinctus (Mydidae: Syllegomydinae) was collected.

Red, vegetated sand dunes (near Klipheuwel, Western Cape) where Afroleptomydas opacicinctus (Mydidae: Syllegomydinae) was collected. Note the small flower in the foreground which was visited by the mydid on a few occasions.

White, vegetated sand dunes at coast (McDougall’s Bay, Northern Cape) where Afroleptomydas consanguinensis (Mydidae: Syllegomydinae) was collected.

White, vegetated sand dunes at coast (McDougall’s Bay, Northern Cape) where Afroleptomydas consanguinensis (Mydidae: Syllegomydinae) was collected.

The habitat in Goegap Nature Reserve (near Springbok, Northern Cape) looked great, but unfortunately we weren’t able to find any Mydidae during our visit.

The habitat in Goegap Nature Reserve (near Springbok, Northern Cape) looked great, but unfortunately we weren’t able to find any Mydidae during our visit.

A Xylocopa carpenter bee on an Aloe species (Putsonderwater, Northern Cape).

A Xylocopa carpenter bee on an Aloe species (Putsonderwater, Northern Cape).

A large centipede (near Nababeep, Northern Cape).

A large centipede (near Nababeep, Northern Cape).

A rhombic egg-eater snake (Dasypeltis scabra) (Droe River, near Vanrhynsdorp, Western Cape).

A rhombic egg-eater snake (Dasypeltis scabra) (Droe River, near Vanrhynsdorp, Western Cape).